Bak-Sneppen backwards

Abstract

We study the backwards Markov chain for the Bak–Sneppen model of biological evolution and derive its corresponding reversibility equations. We show that, in contrast to the forwards Markov chain, the dynamics of the backwards chain explicitly involve the stationary distribution of the model, and from this we derive a functional equation that the stationary distribution must satisfy. We use this functional equation to derive differential equations for the stationary distribution of Bak–Sneppen models in which all but one or all but two of the fitnesses are replaced at each step, subject to certain conditions on the relative locations of the replaced species. This gives a unified way of deriving Schlemm’s expressions for the stationary distributions of the isotropic four-species model, the isotropic five-species model, and the anisotropic three-species model.